Penstemon digitalis-Foxglove Beardtongue, Smooth Beardtongue ETC.

Penstemon digitalis (Foxglove Beardtongue, ETC.) on 6-3-19, #581-12.

Smooth Beardtongue, Foxglove Beardtongue, Foxglove Penstemon

Penstemon digitalis

PEN-stem-on  dig-ee-TAH-liss

Synonyms of Penstemon digitalis (7) (Updated on 3-12-21): Chelone digitalis (Nutt.) Sweet, Penstemon alluviorum Pennell, Penstemon digitalis var. albidus Trautv., Penstemon digitalis f. baueri Steyerm., Penstemon laevigatus subsp. alluviorum (Pennell) R.W.Benn., Penstemon laevigatus subsp. digitalis (Nutt.) R.W.Benn., Penstemon laevigatus var. digitalis (Nutt.) A.Gray

Plants of the World Online by Kew list Penstemon digitalis Nutt. as the correct and accepted scientific name for this species of Penstemon. It was named and described as such by Thomas Nuttall in Transactions of the American Philosophical Society in 1835. However… Penstemon digitalis Nutt. ex Sims, being described as such by John Sims in the Botanical Magazine in 1825.

I sent the editor of Kew an email, like I always do when there is an issue, and he sent a link to the publication. Mr. Sims wrote in 1825 after the description:

“Communicated by our friend Robert Barclay, Esc. to whom the seeds were sent by Professor Nuttall, in March, 1824,under the name we have adopted, thinking is probable, that it may have been published but it in America; otherwise a substantive specific name should be confined to such species as have before constituted a different genus. A hardy perennial. Native of the Arkansa territory…”

Mr. Nuttall wrote The Genera of North American Plants, in two volumes in 1818. I read through both editions and Penstemon digitalis is not mentioned. Afterward, in 1835, Mr. Nuttall wrote about the species in Transactions of the American Philosophical Society, which there were many volumes published. I haven’t read through any of those yet to see what he said. The species had to have been named AND described prior to 1824 for him have sent seeds with the name Penstemon digitalis… 

According to what I have read thus far, Mr. Nuttall (who was an English botanist) was on an expedition from 1818 through 1820 raveling along the Arkansas and Red Rivers. After returning to Philadelphia, he published a book about the expedition in 1821. In 1825 he became the curator of the botanical garden at Harvard. He resigned his position and went on another expedition in 1834 and didn’t return home until 1836… SOOOO…. He wrote the description for Penstemon digitalis in Transactions of the American Philosophical Society in 1835 while he was on an expedition? Hmmm… 

Lewis and Clark and previously collected samples of plants of which many were lost. Some of the specimens Nuttall collected on his expeditions were likely the same as the ones that were previously lost and even possibly named by Meriwether Lewis. The problem is, a name and description have to be accompanied by specimens (in one form or another) to make the name legit.

I also read where Nuttall sent seeds and the description for Penstemon digitalis home to England in 1824, which was later “re-published” by Mr. Sims in Botanical Magazine in 1826, after the seeds grew and the plants flowered (so he could have proper specimens no doubt)… I read where Nuttall “didn’t have time” to publish his description himself until 1835 because his work on “this species” was incomplete… Hmmm… How could he have time?

Sorry, I am just venting because I don’t know the truth of who actually named Penstemon digitalis… I wouldn’t be surprised if Meriwether Lewis actually first named the species and the specimens were lost. Just my opinion. Actually, the Wikipedia article about Mr. Nuttall doesn’t list Penstemon digitalis as a species he named, nor is it on the article about Mr. Sims.

I will probably delete most of what I wrote after I have thought about it for a while… 🙂

POWO is likely to change from Penstemon digitalis Nutt. to Penstemon digitalis Nutt. ex Sims since the latter is from an earlier publication. Sometimes it is true that the early bird gets the worm but not always. I didn’t point out to the editor that other databases have said Penstemon digitalis Nutt. ex Sims for many years… It has been in the IPNI (International Plant Names Index) since at least 2010 when I started writing about plants… 

The genus, Penstemon Schmidel, was named and described as such by Casimir Christoph Schmidel in Icones Plantarum in 1763.

Plants of the World Online lists 276 species in the Penstemon genus (as of 3-26-20 when I am updating this page). The genus us a member of the plant family Plantaginaceae with a total of 105 genera. Those numbers are likely to change.

Distribution map of Penstemon digitalis from Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.plantsoftheworldonline.org/. Retrieved on March 26, 2020.

The above distribution map of Penstemon digitalis is from Plants of the World Online. Areas in green are where the species is native and purple where it has been introduced. The map on the USDA Plants Database for North America is the same. The species may be more widespread than what the maps show. People do buy seed…

THERE ARE SEVERAL LINKS AT THE BOTTOM OF THE PAGE FOR FURTHER READING AND TO HELP WITH A BETTER POSITIVE ID.

Penstemon digitalis (Foxglove Beardtongue, ETC.) on 6-3-19, #581-14.

Penstemon digitalis is a perennial wildflower that attracts several species of insects. I don’t have this species growing on my farm, but I found a few fairly large colonies growing along a highway close to town. It was quite a sight.

Plants grow to 3’ tall, more or less, on stems that are mostly hairless (glabrous). The leaves grow in an alternate pattern along the stems and sometimes in whorls of three. Leaf shape is variable but are mostly lance-shaped.

Penstemon digitalis (Foxglove Beardtongue, ETC.) on 6-3-19, #581-15.

The white flowers are tube-shaped and the corollas are divided into a lower lip with 3 lobes and an upper lip with 2 lobes. The flowers of the colony I was photographing had all white flowers but they can also be splashed with purple. If I had have checked every plant I may have found some with streaks.

I first identified this species as Penstemon tubaeflorus, but a member from iNaturalist suggested they were P. digitalis. The lobes on P. tubaeflorus flowers are closer together than those of the P. digitalis otherwise they are pretty much alike. I am sure there are other characteristics that separate the two species if you want to get down to the nitty-gritty.

Penstemon digitalis (Foxglove Beardtongue) on 5-10-20, #697-40.

While wildflower hunting on a friend’s farm on May 10 in 2020 I ran across this plant that was not in flower yet. I took photos anyway and planned to go back when it was in flower to identify it better. I was unable to go back, but an amazing thing happened later and I was able to identify this plant as Penstemon digitalis.

Penstemon digitalis (Foxglove Beardtongue) on 5-10-20, #697-41.

The Foxglove Beardtongue emerges in the spring with a rosette of long lance-shaped basal leaves. Soon it till start sending up a stem that can grow up to at least 3’ tall. The stems are usually glabrous (hairless) but can also have small hairs toward the tip. Information suggests basal leaves persist even after the stem leaves fall off and the fruit goes to seed in the fall.

Penstemon digitalis (Foxglove Beardtongue) on 5-10-20, #697-42.

Both basal and stem leaves grow in an opposite arrangement. The leaves vary in shape from broadly lanceolate, ovate, obovate, spatulate, and rounded too bluntly or sharply pointed at the tip. The longest leaves can be up to at least 6” long and 2 1/2” wide and may have a slightly reddish tint.

Penstemon digitalis (Foxglove Beardtongue) on 5-10-20, #697-43.

The leaves taper slightly toward the base. I think the basal leaves may have winged petioles whereas the upper leaves are sessile (without petioles), sometimes shallowly cordate (triangular) and clasping (around the stems). Th leaves of the plants I observed and photographed had small teeth along the margins but they can also be smooth. Like I said, the leaves are quite variable…

Penstemon digitalis (Foxglove Beardtongue) on 5-10-20, #697-44.

The above photo shows a good close-up of the stem that has very short hairs. Sometimes the hairs appear to be growing in lines…

Penstemon digitalis (Foxglove Beardtongue) on 6-14-20, #710-38.

On June 14 (2020) I decided to take a walk to the back of the farm even though the grass was very tall for hay. Low and behold, I found a single Penstemon digitalis not far after entering the back section. I have been here since 2013 and this was the first time I had seen one here on this farm. It makes me wonder how it could have got here all of a sudden.

Penstemon digitalis (Foxglove Beardtongue) on 6-14-20, #710-39.

It always amazes me how I can be wildflower hunting and find a single plant of a species and none anywhere else to be found… The above photo shows a flower with the reddish streaks…

Penstemon digitalis (Foxglove Beardtongue) on 6-14-20, #710-40.

Then, as I walked toward the southeast corner of the farm I noticed white flowers from a distance. HOLY COW! There was an entire colony! Until 2017 cows always grazed this lower section. In 2017 and 2018 I cut hay here about July. I don’t normally walk through the hayfield because the grass becomes very thick and tall and it is hard to walk through. Still, it is weird they are here right under my nose and I didn’t know it…

Penstemon digitalis (Foxglove Beardtongue) on 6-14-20, #710-41.

I didn’t have to go anywhere this year besides my own farm to take more photos of this plant but I do need to go back and get some close-ups.

Penstemon digitalis (Foxglove Beardtongue) on 6-14-20, #710-42.

The above photo shows leaves that grow in opposite pairs with small teeth along the margins that are sessile (no petioles) that clasp the stem. Pretty neat, huh?

Penstemon flowers attract a variety of long-tongues bees and butterflies which help with pollination.

I hope to find these again on my farm in 2021 but you just never know… I have been back on the family farm since 2013, like I said, and I had never seen them until 2020… It probably has a lot to do with when the hay was cut. Cows don’t particularly eat these plants, but this area hasn’t been grazed for at least four years.

I have enjoyed photographing and learning about the many wildflowers growing on the farm and other areas. My farm is in Windsor, Missouri in Pettis County (Henry County is across the street and Benton and Johnson aren’t far away). I have grown over 500 different plants and most have pages listed on the right side of the blog. I am not an expert, botanist, or horticulturalist. I just like growing, photographing and writing about my experience. I rely on several websites for ID and a horticulturalist I contact if I cannot figure them out. Wildflowers can be somewhat variable from location to location, so sometimes it gets a bit confusing. If you see I have made an error, please let me know so I can correct what I have written.
I hope you found this page useful and be sure to check the links below for more information. They were written by experts and provide much more information. Some sites may not be up-to-date but they are always a work in progress. If you can, I would appreciate it if you would click on the “Like” below and leave a comment. It helps us bloggers stay motivated. You can also send an email to me at thebelmontrooster@yahoo.com. I would enjoy hearing from you especially if you notice something is a bit whacky.
NOTE: Plants of the World Online is the most up-to-date database. It is very hard for some to keep with name changes these days so you may find a few discrepancies between the websites. Just be patient. Hopefully, someday they will be in harmony. 🙂

FOR FURTHER READING:
PLANTS OF THE WORLD ONLINE (GENUS/SPECIES)
WORLD FLORA ONLINE (GENUS/SPECIES)
WIKIPEDIA (GENUS/SPECIES)
MISSOURI BOTANICAL GARDEN
MISSOURI PLANTS
iNATURALIST
WILDFLOWERSEARCH.ORG
DAVE’S GARDEN
USDA PLANTS DATABASE
ILLINOIS WILDFLOWERS
MINNESOTA WILDFLOWERS
KANSAS WILDFLOWERS AND GRASSES
GO BOTANY
LADY BIRD JOHNSON WILDFLOWER CENTER
FLORA FINDER
FRIENDS OF THE WILDFLOWER GARDEN

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